Carpentaria Becc., Ann. Jard. Bot. Buitenzorg 2: 128 (1885)

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  • Moderate or tall, solitary, unarmed, pleonanthic, monoecious palm. Stem erect, moderate, smooth, grey, ringed with leaf scars. Leaves pinnate, arching or drooping, tips becoming pendulous; sheath forming a prominent crownshaft; petiole very short, deeply channelled adaxially, rounded abaxially, covered with small brown scales; rachis channelled basally to ridged; leaflets abaxially shallowly convex, ± clustered basally, opposite to subopposite distally, long, narrow, lanceolate, tips oblique to truncate, praemorse, often with 2–4 longer prongs, the proximal sometimes tapering to a single or double point, adaxially lightly tomentose, abaxially covered with small brown-centred scales, midrib the only prominent nerve on both surfaces, larger abaxially, transverse veinlets not apparent. Inflorescences infrafoliar, horizontal, appearing rather large, branched to 3 orders basally, to 2–1 orders distally; peduncle very short, stout, flattened, bearing deciduous brown tomentum; prophyll tubular (not seen), caducous; peduncular bract tubular, caducous, shortly beaked, splitting abaxially, adaxially glabrous, abaxially densely to lightly covered in stellate brown scales, scar of 1 incomplete peduncular bract present; rachis much longer than the peduncle, elongate, tapering, bearing many (more than 20) ± angled first-order branches, each subtended by a very small, ridge-like bract; rachillae rather short, slender, spreading, bearing spirally arranged, distant triads of flowers for 2/3 their length and paired to solitary staminate flowers distally; floral bracteoles large, low, rounded. Staminate flowers lateral to the pistillate, symmetrical, ovoid; sepals 3, distinct, broadly imbricate, irregularly rounded, somewhat gibbous; petals 3, distinct, rather broadly ovate, valvate, evenly thickened, adaxially grooved; stamens ca. 33, filaments erect in bud, short, awl-shaped, anthers oblong-elliptical, bifid basally, emarginate apically, dorsifixed near the base, latrorse, connective broad, tanniniferous; pistillode bottle-shaped, as long as the stamens in bud. Pollen ellipsoidal asymmetric; aperture a distal sulcus; ectexine tectate, perforate and micro-chanelled, aperture margin slightly finer than main tectum; infratectum columellate; longest axis ranging from 39–53 µm; post-meiotic tetrads tetrahedral [1/1]. Pistillate flowers ovoid; sepals 3, distinct, imbricate, rounded, margins variously split; petals 3, distinct, broadly ovate and imbricate, tips thick, valvate, opening briefly apically to expose the stigmas at anthesis; staminodes 3, tooth-like, bifid apically; gynoecium asymmetrical, ovoid with a bulge on one side, unilocular, uniovulate, stigmas 3, fleshy, recurved at anthesis, ovule very large, laterally attached, form unknown. Fruit ovoid when fresh, red at maturity, stigmatic remains apical, perianth persistent; epicarp smooth, becoming wrinkled when dry, mesocarp fleshy over a layer of broad, flat fibres anastomosing distally and closely appressed to the endocarp, endocarp thin, glass-like. Seed attached laterally, ovoid, ± pointed, round in cross-section, hilum elongate, raphe branches anastomosing, endosperm homogeneous; embryo basal. Germination adjacent-ligular; eophyll bifid. Cytology: 2n = 32. (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)A

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