Zombia L.H.Bailey, Gentes Herb. 4: 240 (1939)Protologue

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Discussion

Diagnosis

Description

  • Moderate, clustering, spiny, pleonanthic, hermaphroditic palm. Stem erect, slender, covered with persistent, overlapping spiny sheaths, few erect spine-like pneumatophores emerging from the ground near the stem base. Leaves induplicate, palmate, marcescent; sheath expanding into a regular network of fibres, distal fibres reflexed, forming a partial whorl of spines; petiole very slender, elongate, unarmed, semicircular in cross-section; adaxial hastula 3-lobed, center lobe pointed, lateral lobes rounded becoming irregularly tattered, abaxial hastula also pointed with very shallow lateral ridges; blade irregularly divided along the adaxial folds 1/2 – 2/3 to the base into single-fold, lanceolate, rather thin, shortly bifid segments, shiny dark green above, whitish beneath, caducous indumentum along ribs on both surfaces, midrib prominent abaxially, interfold ribs prominent adaxially, transverse veinlets scarcely discernible. Inflorescences interfoliar, shorter than the leaves, branched to 2 orders; peduncle short; prophyll partly included in the subtending the leaf sheath, tubular, 2-keeled, opening and ± 2-lobed distally, longitudinally striate; peduncular bracts lacking; rachis longer than the peduncle, densely tomentose; rachis bracts tubular, longitudinally striate, sparsely tomentose, each with a short pointed lobe, sometimes with short apical splits; first-order branches distant, each bearing a basal, tubular, 2-keeled, 2-lobed, longitudinally striate, sparsely tomentose prophyll; rachillae (second-order branches) sparse, short, spreading, proximal 2 ± subopposite, subtended by narrow bracts, distal rachillae subopposite or alternate, subtending bracts minute, inconspicuous, rachillae slender, glabrous, minutely papillose, bearing rather distant, solitary, subdistichous flowers each subtended by a minute triangular bract. Flowers cream-coloured; perianth shallow, cup-shaped with 6 short, membranous points; stamens 9–12, the filaments short, variable, slender, anthers basifixed, erect, elongate, latrorse; gynoecium obpyriform, unicarpellate, tapered to a large, laterally compressed, cup-like stigma, ovule basal, orthotropous. Pollen ellipsoidal, slight to obvious asymmetry; aperture a distal sulcus; ectexine tectate, perforate, aperture margin ± psilate; infratectum columellate; longest axis 30–38 µm [1/1]. Fruit falling with the perianth attached and stamen bases often obvious, oblong-globose, white, fleshy, large, with apical stigmatic remains; epicarp smooth except for the stigmatic remains, mesocarp fleshy without obvious fibres, endocarp crustaceous. Seed basally attached, very deeply bilobed, the lobes again divided irregularly in a bilobed fashion, the 2 halves connected by a portion in which the embryo lies centrally, endosperm homogeneous within the lobes. Germination and eophyll not recorded. Cytology: 2n = 36. (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)

Anatomy

Relationships

Bibliography

A. J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008
B. World Checklist of Arecaceae