Pinanga Blume, Rumphia 2: 76 (1839)

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Distribution

Map uses TDWG level 3 distributions (http://www.nhm.ac.uk/hosted_sites/tdwg/geogrphy.html)

About 131 species ranging from the Himalayas and south China to New Guinea, with the greatest diversity in the wet areas of the Sunda Shelf; very poorly represented in Papuasia. (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)

Discussion

Diagnosis

Biology And Ecology

Etymology

Uses

Description

  • Very small to robust, solitary or clustered, acaulescent or erect, unarmed, pleonanthic, monoecious palms. Stem very slender to moderate, with elongate or short internodes and conspicuous leaf scars, occasionally stilt-rooted. Leaves undivided and pinnately ribbed, with or without an apical notch, or pinnate; sheaths tubular, forming a well-defined crownshaft, with leaves neatly abscising, very rarely leaves marcescent and crownshaft not well developed; petiole present or absent, adaxially rounded or channelled, abaxially rounded, glabrous or variously indumentose; leaflets 1 to several-fold, regularly to irregularly arranged, acute, acuminate, or lobed, the lobes corresponding to the folds, the apical leaflets almost always lobed, blade occasionally mottled, sometimes paler beneath, often with a wide variety of scales and hairs, transverse veinlets usually obscure. Inflorescence mostly infrafoliar, rarely interfoliar in acaulescent species with marcescent leaves, very rarely bursting through marcescent leaf sheaths (Pinanga simplicifrons), usually rapidly becoming pendulous, occasionally erect, protogynous, unbranched or branching to 1 order only; peduncle usually short, dorsiventrally flattened, glabrous or tomentose; prophyll thin, membranous, 2-keeled, enclosing the inflorescence in bud, quickly splitting to expose the flowers except in P. simplicifrons and P. cleistantha where persistent and enclosing inflorescence up to almost mature fruiting; peduncular bracts absent; rachis usually longer than the peduncle; rachis bracts triangular, usually very inconspicuous; rachillae bearing spiral or distichous triads throughout, or triads in 4 or 6 vertical rows, or, more rarely, spiral proximally and distichous distally; triads sometimes partially sunken in the axis of the rachilla, but well-defined pits not present; floral bracteoles minute. Staminate flowers asymmetrical, sessile, rarely stalked at the base, very rarely the stalk of one flower much longer than the other (P. cleistantha); calyx cupular with 3 triangular, frequently unequal lobes; petals 3, triangular, frequently unequal, joined briefly basally, valvate in bud, much exceeding the calyx lobes, usually very fleshy; stamens rarely 6, usually 12–68, filaments short, anthers linear, latrorse; pistillode absent. Pollen usually ellipsoidal, occasionally oblate triangular, with at least one plane of symmetry, less frequently without symmetry; aperture either a distal sulcus, a distal trichotomosulcus, an extended sulcus or a presumed meridional zonasulcus (rare); ectexine either tectate, semitectate, or intectate; ectexine tectate or semitectate pollen finely to coarsely perforate, finely rugulate-reticulate, finely to coarsely reticulate (in some species the muri perforate or reticulate), discrete, psilate ring-like elements, dense supratectal clavae (in some species vertically striate), or finely reticulate with large smooth, broad-based supratectal spines; ectexine of intectate pollen with semi-coalesced mushroom-like pilae interspersed with dense granulae or spinulae, spines interspersed with dense granulae or small clavae, small and large gemmae interspersed or, urceolae interspersed with small dense clavae; infratectum columellate; longest axis 26–60 µm; post-meiotic tetrads tetragonal, and possibly also tetrahedral [50/128]. Pistillate flowers usually globose, symmetrical, much smaller than the staminate; sepals 3, membranous, striate, imbricate, distinct, or connate proximally with 3 broad, sometimes imbricate lobes distally; petals 3, distinct, imbricate, membranous; staminodes absent; gynoecium unilocular, uniovulate, globose, stigma usually convolute, sessile or on a short style, ovule basally attached, anatropous. Fruiting rachillae usually brightly coloured (reddish or orange). Fruit globose, or ellipsoidal to spindle-shaped, sometimes narrow spindle-shaped and curved (P. salicifolia and others), bright crimson, scarlet, orange or black, very rarely dull brown or green, frequently passing through pink to crimson to black at maturity, stigmatic remains apical; epicarp usually smooth, shiny, with a silky sheen, or dull, mesocarp usually thin, fleshy, sweet, rarely greatly expanding (e.g., P. keahii), endocarp of longitudinal fibres, usually adhering to the seed, becoming free at the basal end only (see Nenga), fruit without a solid beak. Seed conforming to the fruit shape, but usually slightly hollowed at the base, with conspicuous basal hilum and anastomosing raphe branches, endosperm deeply ruminate or, very rarely, subruminate or homogeneous; embryo basal. Germination adjacent-ligular; eophyll bifid or rarely entire with a minute apical cleft. Cytology: 2n = 32. (J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008)

Anatomy

Relationships

Taxonomic accounts

Bibliography

A. J. Dransfield & N. Uhl & C. Asmussen & W.J. Baker & M. Harley & C. Lewis, Genera Palmarum. The evolution and classification of palms. 2008
B. World Checklist of Arecaceae